Award Date

8-1-2024

Degree Type

Dissertation

Degree Name

Doctor of Philosophy (PhD)

Department

Life Sciences

First Committee Member

Brian Hedlund

Second Committee Member

Duane Moser

Third Committee Member

Mira Han

Fourth Committee Member

Jeffery Shen

Fifth Committee Member

Ganqing Jiang

Number of Pages

175

Abstract

High-temperature geothermal springs are ecosystems that resemble early Earth, offering insights into the origins and early evolution of life. In these unique ecosystems, thermophilic microorganisms thrive. However, our knowledge of thermophiles remains limited. A major challenge is that many thermophiles remain uncultivated. The primary goal of this dissertation was to explore the diversity, metabolism, and ecological functions of thermophiles using cultivation-dependent and cultivation-independent approaches. Chapter 1 provides a broad overview of geothermal spring ecosystems and thermophiles, along with a roadmap of the dissertation. Chapter 2 documents a study on the response of specific thermophiles from Gongxiaoshe (GXS) Hot Spring in China to pulses of labile dissolved organic carbon, specifically acetate and aspartate, using quantitative stable isotope probing (qSIP) and metagenomics. The results indicate strong resource partitioning and a wider preference for aspartate incorporation into DNA over acetate. Chapter 3 explores the ecophysiology and evolution of a dominant archaeal species, designated Calditenuis ramacidaminiphagus, from Great Boiling Spring, Nevada, USA. By using fluorescence in situ hybridization coupled with nanoscale secondary ion mass spectrometry (FISH-NanoSIMS), metagenomics, metaproteomics, and laboratory cultures, we demonstrated branched-chain amino acids (BCAAs) specialization in C. ramacidaminiphagus. We also showed that BCAA specialization as a key trait for the genus Calditenuis. Chapter 4 summarizes our research findings and discusses future research perspectives. Chapter 5 describes my contributions to other research projects.

Keywords

extremophiles; geothermal spring; resource partitioning; thermophiles

Disciplines

Biology | Environmental Sciences | Microbiology

File Format

pdf

File Size

11500KB

Degree Grantor

University of Nevada, Las Vegas

Language

English

Rights

IN COPYRIGHT. For more information about this rights statement, please visit http://rightsstatements.org/vocab/InC/1.0/

Available for download on Friday, August 15, 2025


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